Middle Ordovician
Great Ordovician Biodiversity Event
The Middle Ordovician (470-459 Ma) is a time period that occurs after the Floian (Early Ordovician), and before the Sandbian (Late Ordovician)
Geologic Age
470.0–458.4 Ma
Subdivisions
Darriwilian: 467.3±1.1–458.4±0.9 Ma
Dapingian: 470.0±1.4–467.3±1.1 Ma
Eon / Era / Period / Epoch
What happened during this time?
Flora (plants, fungi, bacteria)
The earliest unambiguous indirect fossil evidence of land plants can be traced to Dapingian, which is represented by dyads and tetrahedral tetrads (microfossils) from the Gondwana paleo-continent
Earliest known macrofossils of bryophytes from the late Middle Ordovician of Tennessee (Retallack 2019)
Liverworts, mosses, and hornworts are all present at the site
Plant life existed on the margins of the land in the Dapingian
By the Darriwilian, plants lived along river valleys and flood plains, far from open water
Fossilized fungi hyphae have been recovered from the middle Ordovician of Wisconsin
These were mycorrhizal fungi: symbiotic fungi serve as or are associated with, plants (Redecker et al. 2000)
Retallack (2019) also found endomycorrhiza and lichens
This suggests plants were growing in regions where they were not covered with water and therefore required a method of extracting minerals from primitive soils, at least for substantial periods of time
Fauna
Great Ordovician Biodiversity Event (GOBE)
Explosion of invertebrate life
Evidence of very early vertebrates
Geophysical
Strange impact craters found during this time; Earth may have had a temporary ring system (Tomkin et al. 2024)
Shading of Earth by the ring may have triggered a global icehouse period.