Kungurian Age
The Kungurian (284–272 Ma) is the fourth and last stage of the Early Permian occurring before the Roadian (Middle Permian), and after the Artinskian
Geologic Age
283.5±0.6–272.3±0.5 million years ago
Eon / Era / Period / Epoch
What happened during this time?
Geophysical
The Earth was still in an ice age at the start of this epoch, but glaciers receded around the mid-Permian period as the climate gradually warmed, drying the continent's interiors
Gondwana collided Laurussia and created the Alleghenian orogeny in present-day North America, while the Hercynian orogeny continued in northwestern Europe
These events created the large supercontinent, Pangea, by the middle of this epoch, which would impact the climate
Biological
Flora, spore-bearing
Noeggerathian progymnosperms are present, and dominant in some swamp environments
Extant clubmoss groups, such as the Lycopodiales and Selaginellales are found in the understory of some forests
Marattialean tree ferns dominate along with progymnosperms in the beginning of the Early Permian
Sphenophyllophytes and zygopterid ferns are found in the under-story, and horsetails along rivers and wetlands
True ferns are beginning to appear on the landscape
Flora, seed-bearing
Cycads definitely on the landscape; there is debate on fossils attributed to cycads from the Carboniferous
Cordaites dominate in Euramerica; Glossopterids dominate in Gondwana
Giganopterids were among the most striking and important plants of the Cathaysian flora of Sino-Malaya (Wang 1999)
Seed ferns, such as the Callistophytales and Peltasperms were still on the landscape
Vojnovskyales, a bizarre seed plant group with flower-like cones, appears in the Pennsylvanian, to quickly disappear in the Early Permian
Fauna
Early Permian tillyardembiid insects (Polyneoptera) preserved with pollen on their heads, thoraces, legs and abdomens (Khramov et al. 2023)
This is the earliest finding of pollen-bearing insects,
Judging by the pollen composition, tillyardembiids visited a narrow range of host plants, including Rufloriaceae (Cordaitales)
Tillyardembiids may have been pollen consumers which contributed to pollination, but it is difficult to conclusively determine
"A trophic specialization of this kind could be considered an evolutionary precursor of pollination mutualism"
Additional Resources
Oldest Known Pollen-Carrying Insects (6Mar2023 Smithsonian Magazine; Khramov et al. 2023)